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In strategy I plants (e.g., tobacco), ferric-chelate reductase (FCR) catalyzes the reduction of Fe 3+ to Fe 2+ , which is then assimilated into root cells via the high-affinity metal transporter IRT1 (iron-regulated transporter 1) ( 2+ , Mn 2+ and Cd 2+ ) ( ferritin 1 ( Fer1 ) is induced by iron application in arabidopsis ( FRO2 and FRO8 , two genes encoding ferric-chelate reductase, were induced in roots of both genotypes by Fe deficiency (Figure 5B)